Galapagos Species Checklist

Phalacrocorax harrisi Rothschild, 1898

Cormorán no volador, Galápagos flightless Cormorant

Galápagos flightless Cormorant in Santa Cruz Island, Galápagos. Photo: CDF Archive.
Galápagos flightless Cormorant in Santa Cruz Island, Galápagos. Photo: CDF Archive.










Taxon category: Accepted

Syn.: Nannopterum harrisi (Rothschild, 1898)

Taxon origin

Taxon origin: Endemic




Feeding type: Piscivorous

Trophic role: Carnivorous

Reproduction mode: Exclusively sexual


Distribution: Present on Fernandina and Isabela Island.


  • Wiedenfeld, D.A. (2006) Aves, the Galapagos Islands, Ecuador. Check List 2006 2(2): 1-27.
  • Jiménez-Uzcátegui, G. Milstead, B., Márquez, C., Zabala, J., Buitrón, P., Llerena, A., et al. (2007) Galapagos vertebrates: endangered status and conservation actions. Galapagos Report 2006–2007. Charles Darwin Foundation, Puerto Ayora, p. 104–110.
  • Jiménez-Uzcátegui, G. Wiedenfeld, D.A. (2002) Marine birds. Reserva marina de Galápagos. Línea base de la biodiversidad. Danulat, E. & G.J. Edgar (eds.). Fundación Charles Darwin/Servicio Parque Nacional Galápagos, Santa Cruz, Galápagos, Ecuador. p. 343–372.
  • Granizo, T. Pacheco, C., Rivadeneira, M. B., Guerrero M. & Suárez, L. (eds.) (2002) Libro Rojo de las Aves del Ecuador. SIMBIOE/Conservation International/EcoCiencia/Ministerio del Ambiente/IUCN. Serie Libros Rojos del Ecuador, tomo 2. Quito, Ecuador.
  • Swash, A. Still, R. (2000) Birds, mammals, and reptiles of the Galapagos Islands, an identification guide. Yale University Press, New Haven, Connecticut, USA, 168 pp.
  • Harris, M.P. (1973) The Galápagos avifauna. Condor 75(3): 265-278.
  • Vargas, F.H. (2006) The ecology of small populations of birds in changing climate. Thesis submitted for the degree of Doctor of Philosophy. Lady Margaret Hall, University of Oxoford.
  • Castro, I. Phillips, A. (1996) A Guide to the Birds of the Galapagos Islands. Christopher Helm Publishers Ltd., London.
  • Deem, S.L. Merkel, J., Ballweber, J., Vargas, F.H., Cruz, M.B., & Parker, P.G. (2010) Exposure to Toxoplasma gondii in Galapagos penguins (Spheniscus mendiculus) and flightless cormorants (Phalacrocorax harrisi) in the Galapagos Islands. Journal of Wildlife Diseases 46: 1005-1011.
  • Swarth, H.S. (1931) The Avifauna of the Galapagos Islands. Occ. Pap. Calif. Acad. Sci. 18: 1-299.
  • Travis, E.K. Hernan Vargas, F., Merkel, J., Gottdenker, N., Miller, R.E., & Parker, P.G. (2006) Hematology, plasma chemistry, and serology of the flightless cormorant (Phalacrocorax harrisi) in the Galapagos Islands, Ecuador. Journal of Wildlife Diseases 42(1): 133-141.
  • Bataille, A. Cunningham, A.A., Cedeño, V., Patiño, L., Constantinou, A., Kramer, L.D. & Goodman, S.J. (2009) Natural colonization and adaptation of a mosquito species in Galápagos and its implications for disease threats to endemic wildlife. Proceedings of the National Academy of Sciences of the United States of America 106(25): 10230-10235.
  • Whiteman, N.K. Sánchez, P., Merkel, J., Klompen, H. & Parker, P.G. (2006) Cryptic host specificity of an avian skin mite (Epidermoptidae) vectored by louseflies (Hippoboscidae) associated with two endemic Galápagos bird species. The Journal of Parasitology, 92(6): 1218-1228.
  • Duffie, C.V. Glenn, T.C., Hernan Vargas, F. & Parker, P.G. (2009) Genetic structure within and between island populations of the flightless cormorant (Phalacrocorax harrisi). Molecular Ecology (2009) 18, 2103–2111, doi: 10.1111/j.1365-294X.2009.04179.x
  • Merkel, J. Jones, H.I., Whiteman, N.K., Gottdenker, N., Vargas, H., Travis, E.K., Miller, R.E., & Parker, P.G. (2007) Microfilariae in Galápagos penguins (Spheniscus mendiculus) and flightless cormorants (Phalacrocorax harrisi): genetics, morphology, and prevalence. Journal of Parasitology, 93(3): 495–503.
  • Siers, S. Merkel, J., Bataille, A., Vargas, F.H. & Parker, P.G. (2010) Eccological correlates of microfilariae prevalence in endangered Galápagos birds. Journal of Parasitology, 96(2): 259-272.
  • Parker, P.G. Whiteman, N.K., & Miller, R.E. (2006) Conservation medicine on the Galápagos Islands: partnerships among behavioral, population, and veterinarian scientists. The Auk 123(3): 625-638.
  • Jiménez-Uzcátegui, G. (2010) Monitoreo del pingüino de Galápagos y cormorán no volador 2010. Informe tecnico para la Fundación Charles Darwin y Parque Nacional Galápagos. Puerto Ayora, Ecuador, 18 pp.
  • Kennedy, M. Valle, C.A. & Spencer, H.G. (2009) The phylogenetic position of the Galapagos Cormorant Molecular Phylogenetics and Evolution 53: 94-98.
  • Valle, C.A. (1994) The ecology and evolution of sequential polyandry in Galapagos Cormorants (Compsohalieus Nannopterum harrisi). A Dissertation for the Degree of Doctor of Philosophy. Princeton University. USA, 156 pp.
  • Valle, C.A. (1986) Status of the Galápagos Penguin and Flightless Cormorant Populations in 1985. Noticias Galápagos 43: 16–17.
  • Valle, C.A. Coulter, M.C. (1987) Present status of the Flightless Cormorant, Galápagos Penguin and greater Flamingo populations in the Galápagos Islands, Ecuador, after the 1982–1983 El Niño. Condor 89(2): 276–281.
  • Larrea, C. (2007) Movimiento, dispersión y éxito reproductivo del cormorán no volador Phalacrocorax harrisi en las islas Galápagos. Disertación para obtención de Licenciatura en Ciencias Biológicas. Pontificia Universidad Católica del Ecuador. Quito, Ecuador, 102 pp.
  • Jiménez-Uzcátegui, G. Valle, C.A. & Vargas, F.H. (2012) Longevity records of Flightless Cormorants Phalacrocorax harrisi Marine Ornithology 40: 127–128
  • Carrera-Játiva Rodríguez-Hidalgo, R., Sevilla, C., & G. Jiménez-Uzcátegui. (2014) Gastrointestinal parasites in the Galápagos Penguin Spheniscus mendiculus and the Flightless Cormorant Phalacrocorax harrisi in the Galápagos Islands. Marine Ornithology 42: 77-80.
  • IUCN (2015) The IUCN Red List of Threatened Species. Version 2015-4. <>. Downloaded on 20 November 2015.
  • Freile, J.F. Santander, T., Jiménez-Uzcátegui, G., Carrasco, L., Cisneros-Heredia, D., Guevara, E., Sánchez-Nivicela, M., Tinoco, B. (2019) Lista Roja de las aves del Ecuador Quito, Ecuador. 97 pp.


The mission of the Charles Darwin Foundation and its Research Station is to tackle the greatest threats and challenges to Galapagos through scientific research and conservation action, in order to safeguard one of the world’s most important natural treasures.

Sign up to the latest news

* required

Sign up to the latest news

* required


Av. Charles Darwin s/n, Puerto Ayora, Galápagos, Ecuador
(593) 5 2526-146 / 2527-013 Ext 101
Quito Office
Francisco Andrade Marín
E6-122 y Av. Eloy Alfaro
+593 (2) 2 231 174

The ‘Charles Darwin Foundation for the Galapagos Islands’, in French ‘Fondation Charles Darwin pour les îles Galapagos”, Association internationale sans but lucratif (AISBL), has its registered office at 54 Avenue Louise, 1050 Brussels, Belgium. Trade Registry # 0409.359.103

© 2023 Charles Darwin Foundation. All rights reserved.